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probability to re ect that three of the strands exited to M3 and only one to I2 Delete states have 0 probability since they do not emit any symbols This HMM is then said to model the four sequences, and the probabilities of each sequence can now be determined as the product of all the probabilities along its path For instance, for AGTC: AGTC = 05 097 10 097 025 09 10 097 10 = 0102676 That is, from left to right, 05 represents the link between begin and M1, 097 the probability of matching A in M1, 10 the link between M1 and M2, 097 the match between G and M2, 025 is the link between M2 and I2, 09 the probability of inserting T in I2, 1.

The tree of life is a portrait of evolution based on molecular phylogeny of genes encoding small subunit ribosomal RNA (see Sections 2.7 and 5.5). The three-domain concept provides a framework for re ning hypotheses about how life developed. If developments were based solely upon linear evolutionary trajectories, once a new positive trait arose, it might be expected to remain exclusively within its original line of descent (see Section 5.9). In practice, however, phylogenetic trajectories often depart from linearity. This departure probably is caused by two major factors: (i) genes being transferred laterally between lines of descent; and (ii) uctuating environmental conditions (hence selective pressures) which over the eons foster hereditary discontinuities, via loss of genes and/or their host via extinction. Insights into the robustness of the tree of life can be obtained by examining other traits (morphological, genetic, biochemical) carried by current members of the Bacteria, Archaea, and Eukarya. Table 2.3 provides a comparison of such traits across the three domains of life. The traits listed

includes the work done by the pressure, the macroscopic energy flux, and the transport of heat by conduction 8, = K,AT, . The thermal conductivity of the species is denoted by K, , which is given in SI units [ 5 ] by

0 the link between I2 and M3, 097 the match between C and M3, and nally 10 the link between M3 and end One of the bene ts of an HMM is that the model can be used to generate other sequences not so far explicitly represented For instance, following only the links that have been activated when evaluating the four previous sequences, the new sequence CAGTC is also a sequence of the model if the two insert states are both entered for a sequence: CAGTC = 05 045 05 097 10 097 025 09 10 097 10 = 0023103 One of the problems with HMMs and any probabilistic approach that uses products of probabilities is that resulting probabilities become increasingly smaller with each link or state probability.

7. Select the Create from File radio button to create the OLE object from an existing file. The dialog changes to display new options. (See Figure 6-27.)

In Design view, you lay out a page visually. As your Web page begins to take shape, Design view shows you a close representation of how the page looks when viewed through a browser such as Firefox or Internet Explorer. You can even see active elements, such as QuickTime movies or Shockwave and Flash files in your Web page as you re building it. You can switch to Design view with the View Design menu command or by clicking the Show Design View button on the Document toolbar, described in the section Accessing the Toolbars later in this chapter. As the name suggests, Code view displays the underlying code used to create the document, whether that is HTML, CSS style definitions, or JavaScript whatever code is used to create the page is visible to you in Code view. If you are working in the Coder style workspace, Code view is the default view; but you can also switch to Code view by choosing the View Code menu command or by clicking the Show Code View button on the Document toolbar.